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Actress Melissa McCarthy brought a sugar-buzz sweetness to her turns as likable if occasionally over-excitable young women on series like "Gilmore Girls" (The WB/The CW, 2001-07), "Samantha Who?" (ABC, 2007-09) and "Mike and Molly" (CBS, 2010- ).
A veteran stand-up and improvisational comic, she burst onto the scene as Sookie St. James, best friend to Lorelai Gilmore (Lauren Graham) on "Gilmore," and played variants of the role in subsequent series and feature films like "The Back-Up Plan" (2010) and "Life as We Know It" (2010).

Her starring turn on "Molly" as a an overweight teacher who finds herself in a relationship with a similarly rotund policeman (Billy Gardell) was a fine showcase for her talents, and signaled the launch of her career as leading lady. Meanwhile, she continued to shine in a number of high-profile supporting roles, particularly in the hit female-driven comedy from Judd Apatow, "Bridesmaids" (2011), in which she displayed her exceptional gifts for physical comedy and demonstrated that when it came to scene-stealing, she had few comedic peers.

Born Aug. 26, 1970, in Plainfield, IL, Melissa McCarthy's cousin was Playmate-turned-actress and author Jenny McCarthy, who provide her with her first screen credit in an episode of her short-lived sketch comedy series, "The Jenny McCarthy Show" (MTV, 1997).

After graduating from Joliet Catholic Academy in Joliet, IL, she lit out for New York City, where she made a name for herself on the stand-up comedy circuit. While pursuing this career, she also studied dramatic acting at The Actors Studio, and made regular appearances in theater productions.

McCarthy eventually moved to Los Angeles, where she kept a hand in comedy by performing with the famous Groundlings troupe, while pursuing roles in films and television. A memorable performance as a young woman who enjoyed a gossipy phone relationship with the Lord in John August's film short "God" (1998) led to a supporting role in his indie drama "Go" (1999) which, amusingly enough, also featured her in a phone conversation.

The following year, McCarthy landed her breakthrough role as Sookie on "Gilmore Girls." In the show's original pilot, the role was played by comedian Alex Borstein of "Family Guy" (Fox, 1999-2002; 2005- ), but scheduling conflicts forced her to abandon the role, which lead to McCarthy's casting. As Sookie, McCarthy was able to flex both sides of her acting talent; though Sookie was scatterbrained and occasionally over-excited, she also served as a grounding force for Lauren Graham's Lorelei, especially in regard to her various romantic entanglements. Sookie also provided a healthy role model of the joys of a nuclear family through her relationship with Jackson (Jackson Douglas), her produce provider, whom she would later marry and have children with.

The exposure afforded by the popularity of "Gilmore Girls" led to a string of showy bit parts in major features like "Charlie's Angels" (2000), as well as more substantive roles in indies like "Pumpkin" (2002) and "The Third Wheel" (2002). In the former, she trod a fine line between heartbreak and humor as a bright but lovelorn young woman who is set up on a date with a mentally handicapped man (Hank Harris) by the shallow sorority girl (Christina Ricci) who mentored him, while in the latter, she charmed viewers as Ben Affleck's co-worker, who joined him in monitoring a disastrous date between Luke Wilson and Denise Richards. McCarthy's ebullient voice was also featured in several episodes of the animated series "Kim Possible" (Disney Channel, 2002-07) as DNAmy, a scientist and dedicated collected of stuffed animals who becomes obsessed with genetically altering real creatures to make them "cuter."